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Published in the Worls Rose News, July 2012, pp. 43-54
Authors: Okagi, K.1 , Katsumata, M.1 , Hashimoto, S.1 , Inoue, A.1 , Yomogida, K.2 , Maebara, K.3 , Arihara, H.3 , Tsuji, M.4 , Oyama-Okubo, N.5 , Mikanagi, Y.6
(1 Toyotama International Inc., 2 Yomogida Rose Fragrance Institute Corp., 3 NPO Rose Culture Institute, 4 Hamadera Park Rose Garden, 5 National Agriculture and Food Research Organization - National Institute of Floricultural Science, 6 Natural History Museum and Institute, Chiba